这是一个颇具超现实主义色彩的爱情故事，关于一个不相信真爱的女孩遇上一个疯狂爱上她的男孩：约瑟夫·高登-莱维特饰演的汤姆是以给贺卡撰写各种贺语为生，满脑子充满奇妙浪漫的他不意被自己的女友莎莫（佐伊·丹斯切尔 饰）飞掉。为了反思他和莎莫的问题所在，汤姆于是开始回忆自己和这个女孩过去在一起的500个日子……

Boy meets girl. Boy falls in love. Girl Doesn't. The film tells the story of the relationship between a woman, Summer (Zooey Deschanel), who doesn't believe in true love and a man, Tom (Gordon-Levitt), described as a hopeless romantic, who falls in love with her. Over a span of 500 days, the story moves in a non-linear fashion from the perspective of Tom, who goes from ecstatic giddiness one moment, indulging in a fantasy song and dance sequence at one point, to crippling depression the next. The soundtrack features two songs ('Us' and 'Hero') by Regina Spektor, and 'Sweet Disposition' by Melbourne-based band The Temper Trap. Artists include Black Lips, classic tracks from The Smiths, Doves, Feist, Meaghan Smith and many more.